Ecosystem engineers and their effects have long been part of evolving ecological systems. In evolutionary terms, changes like these brought about by living things, including ecosystem engineers, have been slow and incremental. African elephants convert wooded savanna to open grassland by toppling trees as they browse. Beaver dams alter the flow of rivers, increase dissolved oxygen in downstream waters, create wetlands, and modify streamside zones. Some organisms, like beavers and elephants, change their surroundings so much that they have been called ecosystem engineers. Even unusual or seemingly catastrophic events, like volcanic eruptions, are an integral part of the ecological contexts to which organisms adapt over long time spans.
Those that survive are molded by natural selection as the environment changes. Over millennia, organisms evolve to contend with changes in their environment. All organisms change their environment as they live, grow, and reproduce.
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